Licorice
Extract Consumption and Salivary Testosterone Concentrations
Robert A Josephs, Jennifer
S Guinn, Michelle L Harper, Frederick Askari
Licorice consumption has
been shown to substantially reduce serum testosterone concentration.
An explanation for this result was that the active component in
licorice (glycyrrhizic acid) interfered with 17ß- hydroxysteroid
dehydrogenase, which has been shown in vitro to catalyse the conversion
of androstenedione to testosterone. We
twice attempted to replicate this effect of licorice but could not.
We identified differences between our methods and those of the previous
study and possible statistical anomalies (including inappropriate
use of statistical tests) in the earlier report. Lancet 2001; 358:
1613-14
Armanini and colleagues
(1) reported that serum testosterone concentration was reduced by
35% after 4 days of licorice consumption in seven men. These findings
have important implications for licorice consumers and those who
study testosterone. We were interested in these effects of licorice
and attempted and failed to find an effect of licorice on spatial
cognition (measures of spatial cognition and changes in testosterone
concentration are correlated). (2) Our failure led us to attempt
a replication of Armanini and colleagues' results. We followed Armanini
and colleagues' methods. (1) We gave licorice to men daily for 4
days and measured their testosterone concentrations in saliva. 20
college-aged men were given 5·6 g of licorice (Baschetti
Licorice Extract) daily for 4 days. This preparation
contained the same amount of glycyrrhizic acid used by Armani and
colleagues (0·5 g). Potential participants were excluded if they
indicated current or former use of illicit or controlled substances
or had a history of hypertension. Testosterone
concentration was measured by RIA before the study and on the day
after the study ended. We noted a statistically
insignificant decrease in testosterone
concentration of 9·5% (table).
Hormone concentrations
in college-aged people before and after licorice consumption
Day 0 Day 4 Effect size Power
Study Armanini and colleagues
Serum testosterone 25·7 (7·5) 14·4 (1·5)† 2·51 >0·995 (mmol/L)
Our study (1) Saliva testosterone 0·59 (0·30) 0·53 (0·24) 0·20 0·15
(mmol/L)
Our study (2) Male saliva 0·57 (0·19) 0·53 (0·19) 0·21 0·12 testosterone
(mmol/L) Female saliva 0·27 (0·13) 0·24 (0·18) 0·22 0·13 testosterone
(mmol/L) Male saliva cortisol 60·7 (17·9) 91·0 (21·5)* 1·35 0·90
(mmol/L) Female saliva cortisol 49·7 (17·9) 69·0 (21·5)† 0·99
0·70 (mmol/L) Data are mean (SD). *p=0·001 for day 0 comparison.
†p=0·05 for day 0 comparison.
We repeated the study. We
changed licorice sources (Frutarom; Haifa, Israel), and recruited
a new group of participants (11 college-aged men and ten college-aged
women), and obtained independent validation of glycyrrhizic acid
content (Independent Laboratories; Denver, CO, USA). A further test
of the licorice potency and validity of the experimental procedure
was provided by measuring changes in cortisol concentrations in
response to licorice. Cortisol concentrations increase in response
to licorice consumption because of inhibition of 11ß-hydroxysteroid
dehydrogenase, which catalyses the conversion of cortisol to cortisone.
(3) Concentrations of testosterone and cortisol in saliva were measured
before the study and for the next 4 days (only initial and final
values are presented, table). Testosterone concentration did not
significantly decrease. We compared the results of our two studies
by 2 test. The effects of licorice on testosterone concentration
were the same in both studies (table). We noted statistically significant
increases in cortisol concentrations (table).
We did a meta-analysis to
combine and compare effect sizes: we combined the effect of licorice
on testosterone across our two studies, and compared the size of
this effect with that reported by Armanini and colleagues. The ten-fold
difference in effect size between our results and those of Armanini
and colleagues was statistically significant (p=0·001). We did not
replicate the results of Armanini and colleagues despite substantially
higher statistical power in our studies. The licorice source (Saila)
used by Armanini and colleagues was different to ours. However,
three things suggest that this difference is unimportant. First,
in our second study, we replicated the well-known effect of licorice
on cortisol, and thus we validated the potency of the licorice and
the effectiveness of our experimental procedure. Second, verification
of glycyrrhizic acid content confirmed that our participants ingested
the same quantity of metabolically active component as the participants
in Armanini and colleagues' study. Third, the decrease in testosterone
concentration was the same in both our investigations, despite differences
in licorice products and sample composition. Armanini and colleagues
measured changes in serum testosterone concentrations, whereas we
used concentrations in saliva. Although serum concentration is much
higher than saliva concentration, changes in testosterone concentration
in saliva are equivalent to changes in serum concentration. Saliva
and serum testosterone concentrations are very strongly correlated.
(4) Therefore, this procedural difference does not explain the large
discrepancy between our results and those of Armanini and colleagues.
Without examination of Armanini and colleagues' raw data, we can
only speculate as to why our results are so different from theirs.
However, Armanini and colleagues' statistical analysis could provide
an explanation for their effect. They reported a drop in SD of 80%
from the start to the end of their study (table). This reduction
in error variance violates assumptions that underly the statistical
tests that they used and renders them inappropriate. Furthermore,
Armanini and colleagues' large pre-administration error variance
indicates that at least one of their participants probably began
the study with a testosterone concentration that was either substantially
greater or lower than the mean. The effect sizes suggest that this
participant had a higher concentration of testosterone than the
mean. If this assumption is correct, then the remaining participants
began the study with testosterone concentrations that were below
the mean. For these participants, the actual licorice-induced decrease
in testosterone concentration was more modest than the mean. Therefore,
removal of statistical outliers (standard procedure is to remove
participants whose testosterone concentrations are greater than
2 SD than the mean) would lower Armanini and colleagues' effect
to be closer to ours. Armanini and colleagues
concluded that men with low libido or hypertension should be questioned
about licorice ingestion. We confirm that licorice increases cortisol,
suggesting a contraindication for its use in people with hypertension.
Because we detected only a small reduction in testosterone, we cannot
confirm that people with low libido should avoid licorice consumption,
nor can we strongly endorse licorice for use in behavioural or medical
research on testosterone.
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Palermo M. Reduction of serum testosterone in men by liquorice.
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Cognitive pattern in men and women is influenced by fluctuations
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3. Heilmann P, Heide J, Hundertmark
S, Schoneshofer M. Administration of glycyrrhetinic acid: significant
correlation between serum levels and the cortisol/cortisone-ratio
in serum and urine. Exp Clin Endocrinol Diabetes 1999; 107: 370-78.
[PubMed]
4. Vittek J, L' Hommedieu
D, Gordon G, Rappaport S, Southren A. Direct radioimmunoassay (RIA)
of salivary testosterone: correlations with free and total serum
testosterone. Life Sci 1985; 37: 711-16. [PubMed]
Department of Psychology,
University of Texas at Austin, TX, 78712, USA (R A Josephs PhD,
J S Guinn BS, M L Harper MS); and Department of Internal Medicine,
University of Michigan, Ann Arbor, MI (F Askari PhD)